- ↑ 1.0 1.1 Masaki Miya et al., "Evolutionary Origin of the Scombridae (Tunas and Mackerels): Members of a Paleogene Adaptive Radiation with 14 Other Pelagic Fish Families" in PLOS One (4 Septembris 2013) fig. 7, tab. S4-S5, textus S1
- ↑ E. Dartevelle, E. Casier, "Les poissons fossiles du Bas-Congo et des régions voisines" in Annales du Musée Royal du Congo Belge A ser. 3 vol. 2 (1959) pp. 257–568; C. Arambourg, J. Signeux, "Les vertébrés fossiles des gisements de phosphates (Maroc-Algérie-Tunisie)" in Notes et mémoires du Service Géologique du Maroc vol. 92 (1952) pp. 1–372; K. A. Monsch, "Revision of the scombroid fishes from the Cenozoic of England" in Transactions of the Royal Society of Edinburgh: Earth Sciences vol. 95 (2004) pp. 445–489; R. Zorzin et al., "Il giacimento a pesci e piante fossili dell'Eocene inferiore di Monte Solane (Verona)" in Bollettino del Museo Civico di Storia Naturale di Verona: Geologia Paleontologia Preistoria vol. 35 (2011) pp. 57–64, cf. Luca Giusberti et al., "A new Fossil-Lagerstätte from the Lower Eocene of Lessini Mountains(northern Italy): a multidisciplinary approach" in Palaeogeography, Palaeoclimatology, Palaeoecology vol. 403 (2014) pp. 1-15
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