| fossil_range = <br>{{Fossil range|390|0|r=[[Devoniano MedioDevonium]]– medium > [[Holocaenum|PraesensRecens]],<ref name=Grzegorz>{{Cite journal|author=Grzegorz Niedźwiedzki, Piotr Szrek, Katarzyna Narkiewicz, Marek Narkiewicz, Per E. Ahlberg |title=Tetrapod trackways from the early Middle Devonian period of Poland |journal=Nature |volume=463 |pages=43–8 |year=2010 |doi=10.1038/nature08623|pmid=20054388|issue=7277|bibcode = 2010Natur.463...43N}}</ref> {{Fossil range|390|0}}