- ↑ 1.0 1.1 Masaki Miya et al., "Evolutionary Origin of the Scombridae (Tunas and Mackerels): Members of a Paleogene Adaptive Radiation with 14 Other Pelagic Fish Families" in PLOS One (4 Septembris 2013) fig. 7, tab. S4-S5, textus S1
- ↑ A. F. Bannikov, "A new genus and species of putative centrolophid fish (Perciformes, Stromateoidei) from the Eocene of Bolca, northern Italy" in Bollettino del Museo Civico di Storia Naturale di Verona: Geologia Paleontologia Preistoria vol. 24 (2000) pp. 37–46; C. A. Papazzoni, E. Trevisani, "Facies analysis, palaeoenvironmental reconstruction, and biostratigraphy of the Pesciara di Bolca (Verona, northern Italy): an early Eocene Fossil-Lagerstätte" in Palaeogeography, Palaeoclimatology, Palaeoecology vol. 242 (2006) pp. 21-35